Labyrinthulomycetes The Labyrinthulomycetes or Labyrinthulea are a class of protists that produce a network of or tubes, which serve as tracks for the cells to glide along and absorb nutrients for them. The two main groups are the labyrinthulids and thraustochytrids . They are mostly marine , commonly found as parasites on algae and seagrasses or as decomposers on dead plant material . They also include some parasites of marine invertebrates .Characteristics Although they are outside the cells, the filaments of Labyrinthulomycetes are surrounded by a membrane . They are formed and connected with the cytoplasm by a unique organelle called a sagenogen or bothrosome. The cells are uninucleated and typically ovoid, and move back and forth along the amorphous network at speeds varying from 5-150 μm per minute. Among the labyrinthulids, the cells are enclosed within the tubes , and among the thraustochytrids, they are attached to their sides.Classification Labyrinthulomycetes/Labyrinthulea used to belong to the defunct fungal phylum Labyrinthulomycota. They were originally considered unusual slime moulds , although they are not very similar to the other sorts. The structure of their zoospores and genetic studies show them to be a primitive group of heterokonts, but their classification and treatment remains somewhat unsettled. This class has usually two orders, Labyrinthulales and Thraustochytriales, or Labyrinthulida and Thraustochytrida , but a third has recently been proposed. Genus Stellarchytrium FioRito & Leander 2016 Family Oblongichytriidae Cavalier-Smith 2012 * Genus Oblongichytrium Yokoyama & Honda 2007 Order Labyrinthulida Doflein 1901 * Family Aplanochytriidae Leander ex Cavalier-Smith 2012 ** Genus Aplanochytrium Bahnweg & Sparrow 1972 * Family Labyrinthulidae Cienkowski 1867 ** Genus Labyrinthomyxa Duboscq 1921 ** Genus Pseudoplasmodium Molisch 1925 ** Genus Labyrinthula Cienkowski 1864 Order Amphitremida Gomaa et al . 2013 * Family Diplophryidae Cavalier-Smith 2012 ** Genus Diplophrys Barker 1868 * Family Amphitrematidae Poche 1913 ** Genus Paramphitrema Valkanov 1970 ** Genus Archerella Loeblich & Tappan 1961 ** Genus Amphitrema Archer 1867 Order Thraustochytriida Alderman & 1974 * Genus Pyrrhosorus Juel 1901 * Genus Thanatostrea Franc & Arvy 1969 * Family Althorniidae Cavalier-Smith 2012 ** Genus Althornia Jones & Alderman 1972 * Family Sorodiplophryidae Cavalier-Smith 2012 ** Genus Sorodiplophrys Olive & Dykstra 1975 * Family Amphifilidae Cavalier-Smith 2012 ** Genus Amphifila Cavalier-Smith 2012 ** Genus Fibrophrys columna Takahashi et al. 2016 * Family Thraustochytriidae Sparrow ex Cejp 1959 ** Genus Japonochytrium Kobayasi & Ôkubo 1953 ** Genus Monorhizochytrium Doi & Honda 2017 ** Genus Sicyoidochytrium Yokoy., Salleh & Honda 2007 ** Genus Aurantiochytrium Yokoy. & Honda 2007 ** Genus Ulkenia Gaertn. 1977 ** Genus Parietichytrium Yokoy., Salleh & Honda 2007 ** Genus Botryochytrium Yokoy., Salleh & Honda 2007 ** Genus Schizochytrium Goldst. & Belsky emend. Booth & Mill. ** Genus Thraustochytrium Sparrow 1936 ** Genus Hondaea Amato & Cagnac 2018 ** Genus Labyrinthulochytrium Hassett & Gradinger 2018Genetic code The labyrinthulomycete Thraustochytrium aureum is notable for the alternative genetic code of its mitochondria which use TTA as a stop codon instead of coding for Leucine . This code is represented by NCBI translation table 23, Thraustochytrium mitochondrial code.Gallery
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