Kauffman–White classification
The Kauffmann–White classification or Kauffmann and White classification scheme is a system that classifies the genus Salmonella into serotypes, based on surface antigens. It is named after Philip Bruce White and Fritz Kauffmann. First the "O" antigen type is determined based on oligosaccharides associated with lipopolysaccharide. Then the "H" antigen is determined based on flagellar proteins. Since Salmonella typically exhibit phase variation between two motile phenotypes, different "H" antigens may be expressed. Salmonella that can express only one "H" antigen phase consequently have motile and non-motile phenotypes and are termed monophasic, whilst isolates that lack any "H" antigen expression are termed non-motile. Pathogenic strains of Salmonella Typhi, Salmonella Paratyphi C, and Salmonella Dublin carry the capsular "Vi" antigen, which is a special subtype of the capsule's K antigen.
Kauffmann–White classification for ''Salmonella''
;Salmonella serotype : :;Examples:
Salmonella enterica serotype Typhimurium 1,4,5,12:i:1,2
monophasic variant of Salmonella Typhimurium 1,4,5,12:i:-
"O"-group | Serovar | "O" antigens | Phase 1 "H" antigens | Phase 2 "H" antigens |
A | S.Paratyphi A | 1,2,12 | a | no phase 2 antigen |
S. Paratyphi A var. Durazzo | 2,12 | a | no phase 2 antigen | |
B | S. Paratyphi B | 1,4,5,12 | b | 1,2 |
S. Paratyphi B var. Odense | 1,4,12 | b | 1,2 | |
S. Java | 1,4,5,12 | b | ||
S. Limete | 1,4,12,27 | b | 1,5 | |
S. Typhimurium | 1,4,5,12 | i | 1,2 | |
S. Typhimurium var. Copenhagen | 1,4,12 | i | 1,2 | |
S. Agama | 4,12 | i | 1,6 | |
S. Abortus-equi | 4,12 | no phase 1 antigen | e,n,x | |
S. Abortus-ovis | 4,12 | c | 1,6 | |
S. Agona | 4,12 | f,g,s | no phase 2 antigen | |
S. Brandenburg | 4,12 | l,v | e,n,z15 | |
S. Bredeney | 1,4,12,27 | l,v | 1,7 | |
S. Derby | 1,4,5,12 | f,g | no phase 2 antigen | |
S. Heidelberg | 1,4,5,12 | r | 1,2 | |
S. Saintpaul | 1,4,5,12 | e,h | 1,2 | |
S. Salinatis | 4,12 | d,e,h | d,e,n,z15 | |
S. Stanley | 4,5,12 | d | 1,2 | |
C1 | S. Paratyphi C | 6,7, | c | 1,5 |
S. Choleraesuis | 6,7 | c | 1,5 | |
S. Choleraesuis var. Kunzendorf | 6,7 | 1,5 | ||
S. Decatur | 6,7 | c | 1,5 | |
S. Typhisuis | 6,7 | c | 1,5 | |
S. Bareilly | 6,7 | y | 1,5 | |
S. Infantis | 6,7 | r | 1,5 | |
S. Menston | 6,7 | g,s,t | no phase 2 antigen | |
S. Montevideo | 6,7 | g,m,s | no phase 2 antigen | |
S. Oranienburg | 6,7 | m,t | no phase 2 antigen | |
S. Thompson | 6,7 | k | 1,5 | |
C2 | S. Bovismorbificans | 6,8 | r | 1,5 |
S. Newport | 6,8 | e,h | 1,2 | |
D | S. Typhi | 9,12,Vi | d | no phase 2 antigen |
S. Ndolo | 9,12 | d | 1,5 | |
S. Dublin | 1,9,12,Vi | g,p | no phase 2 antigen | |
S. Enteritidis | 1,9,12 | g,m | no phase 2 antigen | |
S. Gallinarum | 1,9,12 | no phase 1 antigen | no phase 2 antigen | |
S. Pullorum | ,9,12 | no phase 1 antigen | no phase 2 antigen | |
S. Panama | 1,9,12 | l,v | 1,5 | |
S. Miami | 1,9,12 | a | 1,5 | |
S. Sendai | 1,9,12 | a | 1,5 | |
E1 | S. Anatum | 3,10 | e,h | 1,6 |
S. Give | 3,10 | l,v | 1,7 | |
S. London | 3,10 | l,v | 1,6 | |
S. Meleagridis | 3,10 | e,h | l,w | |
E2 | S. Cambridge | 3,15 | e,h | l,w |
S. Newington | 3,15 | e,h | 1,6 | |
E3 | S. Minneapolis | ,,34 | e,h | 1,6 |
E4 | S. Senftenberg | 1,3,19 | g,s,t | no phase 2 antigen |
S. Simsbury | 1,3,19 | no phase 1 antigen | z27 | |
F | S. Aberdeen | 11 | i | 1,2 |
G | S. Cubana | 1,13,23 | z29 | no phase 2 antigen |
S. Poona | 13,22 | z | 1,6 | |
H | S. Heves | 6,14,24 | d | 1,5 |
S. Onderstepoort | 1,6,14,25 | e,h | 1,5 | |
I | S. Brazil | 16 | a | 1,5 |
S. Hvittingfoss | 16 | b | e,n,x | |
Others | S. Kirkee | 17 | b | 1,2 |
S. Adelaide | 35 | f,g | no phase 2 antigen | |
S. Locarno | 57 | z29 | z42 |
- Antigens in brackets are those that are rarely expressed in that serovar.
Representative stock of antisera
A common set of working antisera is shown below:O-antisera | H-antisera |
polyvalent-O, groups A-G | polyvalent-H, specific and non-specific |
2-O, group A | polyvalent-H, non-specific factors 1,2,5,6,7 |
4-O, group B | a-H |
6, 7-O, group C1 | b-H |
8-O, group C2 | c-H |
9-O, group D | d-H |
3, 10, 15, 19-O group E | e,h-H |
11-O, group F | f,g-H |
13, 22-O, group G | g,m-H |
i-H | |
k-H | |
l,v-H | |
m,t-H | |
r-H |
Laboratories that are likely to investigate typhoid also carry antiserum raised against the Vi antigen.
A set of "Rapid Diagnostic Sera" is also held and is used for determination of common specific H-antigens except i-H. After obtaining a positive agglutination with the polyvalent-H specific and non-specific antiserum, the three RDS antisera are used to identify the H antigen present. Depending on the pattern of positive and negative reactions with the RDS antisera, the specific H antigen may be identified:
antigen | RDS1 | RDS2 | RDS3 |
b | agglutination | agglutination | no agglutination |
d | agglutination | no agglutination | agglutination |
E | agglutination | agglutination | agglutination |
G | no agglutination | no agglutination | agglutination |
k | no agglutination | agglutination | agglutination |
L | no agglutination | agglutination | no agglutination |
r | agglutination | no agglutination | no agglutination |
E = polyvalent for eh, enx, etc.
G = polyvalent for gm, gp, etc.
L = polyvalent for lv, lw, etc.