West Branch Run begins in Jordan Township, Lycoming County and flows southwest for more than a mile. It then turns south-southwest and flows for slightly over a mile until it exits Jordan Township. Upon leaving Jordan Township, the stream enters Pine Township. Here, it turns southwest for a short distance before turning southeast near Pine Summit. The stream then flows through a fairly narrow valley known as California Hollow for some distance. At the end of the valley, it receives Shingle Run, its only named tributary, and abruptly turns south. After more than a mile, West Branch Run passes the ShoemakerCovered Bridge, crossing Pennsylvania Route 442on the way, and turns southeast. It then crosses Pennsylvania Route 442 again and shortly afterwards reaches its confluence with Little Fishing Creek. West Branch Run joins Little Fishing Creek upstream of its mouth.
Tributaries
The only named tributary of West Branch Run is Shingle Run. Shingle Run joins West Branch Run upstream of its mouth. Its watershed has an area of.
Hydrology, geography and geology
The elevation of West Branch Run near its mouth is above sea level. At its headwaters, the stream has an elevation of approximately above sea level. West Branch Run flows through a valley known as California Hollow in Pine Township, Columbia County. West Branch Run has high levels of dissolved oxygen.
Watershed
The watershed of West Branch Run has an area of. There are no roads along West Branch Run where it flows through California Hollow. The stream is one of only a few streams in Columbia County with this property. The Shoemaker Covered Bridge was built over West Branch Run in 1884 by T.S. Christian. The bridge is long. It was rehabilitated in 2008. A bridge carrying the road T-720 was built over the stream in 1936. It is long.
Biology
California Hollow, which West Branch Run flows through, is listed as a "locally significant site" on the Columbia County Natural Areas Inventory. Hemlock and white pine inhabit the slopes of California Hollow. According to the Columbia County Natural Areas Inventory, it may have a high level of diversity among native plants and few invasive plants. Trout reproduce naturally in West Branch Run.