Wentletrap Wentletraps are small, often white, very high-spired, predatory or ectoparasitic sea snails , marine gastropod mollusks in the family Epitoniidae . The word wentletrap originated in Dutch, and it means spiral staircase . These snails are sometimes also called "staircase shells", and "ladder shells". The family Epitoniidae belongs to the superfamily Epitonioidea . Since 2017 this family also includes the former families Janthinidae and Nystiellidae , all part of the informal group Ptenoglossa . Epitoniidae is a rather large family , with an estimated number of species about 630.Distribution Wentletraps inhabit all seas and oceans worldwide , from the tropical zones to the Arctic and Antarctic zones.Most species of wentletrap are white, and have a porcelain-like appearance. They are notable for their intricately geometric shell architecture, and the shells of the larger species are prized by collectors. The more or less turret-shaped shell consists of tightly-wound, convex whorls, which create a high, conical spiral . Fine or microscopic spiral sculpture is present in many species. The shells sometimes feature an umbilicus . Wentletrap shells have a roundish or oval aperture , but its inner lip is often reduced to strip of callus. The round and horny operculum is and fits the aperture tightly . Most of the species in the family are small to minute, although some are larger, and overall the adult shell length in the family varies between 0.6 and 11.7 cm. Within the genus Epitonium , the type genus of the family, the shell has predominantly axial sculpture of high, sharply ribbed "costae". These costae may offer some protection against other predatory snails, which would find it difficult or impossible to bore a hole in a shell with such obstructions.Ecology Wentletraps are usually found on sandy bottoms near sea anemones or corals, which serve as a food source for them. Some species are foragers and search for anemones. Little is known about the biology of most wentletraps. Keen has his literature most cited. He observed that many wentletraps reveal a hint of purple body color , suggestive of carnivorous feeding. The animal can exude through its salivary gland a pink or purplish dye that may have an anaesthetic effect on its prey. Keen also cited direct observation of a wentletrap feeding by insertion of its proboscis into a sea anemone . A sequence of a wentletrap feeding on an anemone has been published. These snails also prey on corals and other coelenterates. Female wentletraps lay egg capsules that are bound together with a supple string. The young emerge from these capsules as free-swimming larvae.Genera Genera within the family Epitoniidae include: Acirsa Mörch , 1857 Acrilloscala Sacco 1891 Alexania Strand, 1928 Alora Amaea H. & A. Adams, 1853 Boreoscala Kobelt, 1902 Cerithiscala de Boury, 1887 Chuniscala Thiele, 1928 Cingulacirsa Higo & Goto, 1993 Cirsotrema Mörch, 1852 Claviscala de Boury, 1909 Couthouyella Bartsch 1909 Crebriscala de Boury, 1909 Cycloscala Dall, 1889 Cylindriscala de Boury, 1909 Depressiscala de Boury 1909 Ecclesiogyra Dall, 1892 Eglisia Gray, 1842 Epidendrium A. Gittenberger & E. Gittenberger, 2005 Epifungium A. Gittenberger & E. Gittenberger, 2005 Epitonium Röding , 1798 Filiscala de Boury, 1911 Foratiscala de Boury 1887 Funiscala de Boury, 1890 Globiscala de Boury, 1909 Gregorioiscala Cossman, 1912 Gyroscala de Boury, 1887 Janthina Röding, 1798 Kurodacirsa Masahito & Habe, 1975 Minabescala Nakayama, 1994 Narvaliscala Iredale , 1936 Opalia H. & A. Adams, 1853 Periapta Bouchet & Waren, 1986 Plastiscala Iredale, 1936 Problitora Iredale, 1931 Punctiscala Philippi , 1844 Recluzia Petit de la Saussaye , 1853 Rutelliscala Kilburn, 1985 Sthenorhytis Conrad 1862 Surrepifungium A. Gittenberger & E. Gittenberger, 2005 † Turriscala de Boury, 1890 † Variciscala de Boury, 1909 Varicopalia Kuroda MS, 1960 ;Genera brought into synonymy Acrilla H. Adams, 1860: synonym of Amaea H. Adams & A. Adams, 1853 Acutiscala de Boury, 1909 : synonym of Epitonium Röding, 1798 Amiciscala Jousseaume 1912 : synonym of Epitonium Röding, 1798 Asperiscala de Boury, 1909: synonym of Epitonium Röding, 1798 Cinctiscala de Boury 1909 : synonym of Asperiscala de Boury, 1909 Cirratiscala de Boury, 1909 : synonym of Epitonium Röding, 1798 Clathroscala de Boury 1889 : synonym of Amaea H. Adams & A. Adams, 1853 Clathrus Oken 1815 : synonym of Epitonium Röding, 1798 Compressiscala Masahito & Habe 1976 : synonym of Gregorioiscala Cossmann, 1912 Dannevigena Iredale 1936 : synonym of Cirsotrema Mörch, 1852 Foliaceiscala de Boury 1912 : synonym of Epitonium Röding, 1798 Fragiliscala Azuma 1962 : synonym of Amaea H. Adams & A. Adams, 1853 Fragilopalia Azuma 1972 : synonym of Amaea H. Adams & A. Adams, 1853 Glabriscala de Boury 1909 : synonym of Epitonium Röding, 1798 Lampropalia Kuroda & Ito, 1961 : synonym of Cylindriscala de Boury, 1909 Mazescala Iredale 1936 : synonym of Epitonium Röding, 1798 Nipponoscala Masahito & Habe 1973 : synonym of Epitonium Röding, 1798 Nodiscala de Boury 1889 : synonym of Opalia H. Adams & A. Adams, 1853 Nystiella Clench & Turner, 1952 : synonym of Opaliopsis Thiele, 1928 Sagamiscala Masahito, Kuroda & Habe, 1971 : synonym of Globiscala de Boury, 1909 Scala Mörch, 1852 : synonym of Epitonium Röding, 1798 Scalina Conrad, 1865 : synonym of Amaea H. Adams & A. Adams, 1853 Spiniscala de Boury, 1909 : synonym of Epitonium Röding, 1798 Turbiniscala de Boury 1909 : synonym of Epitonium Röding, 1798 Viciniscala de Boury 1909 : synonym of Epitonium Röding, 1798
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