Sphaeropteris Sphaeropteris is a genus of tree fern in the family Cyatheaceae . It has been treated as a subgenus within the genus Cyathea , but is accepted in the Pteridophyte Phylogeny Group classification of 2016.Description Species of Sphaeropteris have a treelike habit, usually with a single tall stem, and large fronds to or more in length. The stalk of the frond is strawlike or purple in colour, with pale to brown scales. The sori are rounded, with or without indusia .Sphaeropteris is now separated from the other genera in the family Cyatheaceae primarily on the basis of molecular phylogenetic studies. However, the scales on the stalks provide a morphological distinction. Sphaeropteris has scales without distinct margins, whereas the other genera have scales with distinct margins.Taxonomy The genus Sphaeropteris was erected by Johann Jakob Bernhardi in 1801. It is placed in the family Cyatheaceae. The division of the family into genera has had a long and controversial history. Three or four clades have been suggested based on molecular phylogenetic studies. The Pteridophyte Phylogeny Group classification of 2016 accepts three genera, related as shown in the cladogram below. Sphaeropteris is sister to the remaining members of the family. Older sources , such as the New Zealand Organisms Register as of 2000, place Sphaeropteris within a very broadly defined Cyathea .Species , the Checklist of Ferns and Lycophytes of the World accepted the following species:Sphaeropteris aciculosa R.M.TryonSphaeropteris aeneifolia R.M.TryonSphaeropteris agatheti R.M.TryonSphaeropteris albidosquamata R.M.TryonSphaeropteris albifrons R.M.TryonSphaeropteris albosetacea R.M.TryonSphaeropteris angustipinna R.M.TryonSphaeropteris aramaganensis R.M.TryonSphaeropteris arthropoda R.M.TryonSphaeropteris assimilis R.M.TryonSphaeropteris atrospinosa R.M.TryonSphaeropteris atrox R.M.TryonSphaeropteris auriculifera R.M.TryonSphaeropteris binuangensis R.M.TryonSphaeropteris brackenridgei R.M.TryonSphaeropteris brunei R. TryonSphaeropteris brunoniana R.M.TryonSphaeropteris capitata R.M.TryonSphaeropteris carrii R.M.TryonSphaeropteris cooperi R.M.TryonSphaeropteris cuatrecasasii R. TryonSphaeropteris curranii R.M.TryonSphaeropteris discophora R.M.TryonSphaeropteris elliptica R.M.TryonSphaeropteris elmeri R.M.TryonSphaeropteris excelsa R.M.TryonSphaeropteris feani R.M.TryonSphaeropteris felina Pic. Serm.Sphaeropteris fugax R.M.TryonSphaeropteris fusca R.M.TryonSphaeropteris gardneri R.M.TryonSphaeropteris glauca R.M.TryonSphaeropteris horrida R. TryonSphaeropteris inaequalis R.M.TryonSphaeropteris insignis R. TryonSphaeropteris insularum R.M.TryonSphaeropteris integra R.M.TryonSphaeropteris intermedia R.M.TryonSphaeropteris intramarginalis WindischSphaeropteris ledermannii R.M.TryonSphaeropteris leucolepis R.M.TryonSphaeropteris leucotricha R.M.TryonSphaeropteris lockwoodiana P.G.WindischSphaeropteris lunulata R.M.TryonSphaeropteris macarenensis R. TryonSphaeropteris magna R.M.TryonSphaeropteris marginata R.M.TryonSphaeropteris medullaris Bernh.Sphaeropteris megalosora R.M.TryonSphaeropteris mertensiana R.M.TryonSphaeropteris microlepidota R.M.TryonSphaeropteris mollicula R. TryonSphaeropteris moluccana R.M.TryonSphaeropteris moseleyi R.M.TryonSphaeropteris myosuroides R. TryonSphaeropteris nigricans R.M.TryonSphaeropteris novae-caledoniae R.M.TryonSphaeropteris obliqua R.M.TryonSphaeropteris obscura R.M.TryonSphaeropteris papuana R.M.TryonSphaeropteris parianensis P.G.WindischSphaeropteris parksiae R.M.TryonSphaeropteris parvifolia R.M.TryonSphaeropteris parvipinna R.M.TryonSphaeropteris persquamulifera R.M.TryonSphaeropteris philippinensis R.M.TryonSphaeropteris pilulifera R.M.TryonSphaeropteris polypoda R.M.TryonSphaeropteris procera R.M.TryonSphaeropteris propinqua R.M.TryonSphaeropteris pukuana Lehnert & CoriticoSphaeropteris pulcherrima R.M.TryonSphaeropteris quindiuensis R. TryonSphaeropteris robinsonii R.M.TryonSphaeropteris robusta R.M.TryonSphaeropteris rosenstockii R.M.TryonSphaeropteris runensis R.M.TryonSphaeropteris sarasinorum R.M.TryonSphaeropteris senex R.M.TryonSphaeropteris senilis R. TryonSphaeropteris setifera R.M.TryonSphaeropteris sibuyanensis R.M.TryonSphaeropteris squamulata R.M.TryonSphaeropteris stipitipinnula R.M.TryonSphaeropteris strigosa R.M.TryonSphaeropteris subsessilis R.M.TryonSphaeropteris suluensis R.M.TryonSphaeropteris tenggerensis R.M.TryonSphaeropteris teysmannii R.M.TryonSphaeropteris tomentosa R.M.TryonSphaeropteris tomentosissima R.M.TryonSphaeropteris trichodesma R.M.TryonSphaeropteris trichophora R.M.TryonSphaeropteris tripinnata R.M.TryonSphaeropteris tripinnatifida R.M.TryonSphaeropteris truncata R.M.TryonSphaeropteris vaupelii R.M.TryonSphaeropteris verrucosa R.M.TryonSphaeropteris vittata R.M.TryonSphaeropteris wallacei R.M.TryonSphaeropteris werneri R.M.TryonSphaeropteris womersleyi R.M.TryonSphaeropteris zamboangana R.M.TryonDistribution and habitat The native distribution of the genus Sphaeropteris extends from southern China down through eastern tropical Asia to New Zealand , with a separate area in Central America and north-western South America . Some species are also naturalized in Africa and western Australia . Species of Sphaeropteris are large plants and require space to grow. They usually prefer less shaded conditions than species of Alsophila . They are found in rain forests and tropical montane forests , often in clearings or on the margins, from the canopy layer down to the understorey layer. They are also found in ravines, swamps and disturbed areas.
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