Sallie W. Chisholm Sallie Watson "Penny" Chisholm is a U.S. biological oceanographer at the Massachusetts Institute of Technology . She is an expert in the ecology and evolution of ocean microbes .Early life and education Chisholm was born in Marquette, Michigan and graduated from Marquette Senior High School in 1965. She attended Skidmore College and earned a Ph.D. from SUNY Albany in 1974.Career Chisholm has been a faculty member at the Massachusetts Institute of Technology since 1976. Her research has focused on the ecology of marine phytoplankton . Chisholm's early work focused on the processes by which such plankton take up nutrients and the manner in which this affects their life cycle on diurnal time scales . This led her to begin using flow cytometry which can be used to measure the properties of individual cells. The application of flow cytometry to environmental samples led Chisholm and her collaborators to the discovery that small plankton accounted for a much more substantial part of marine productivity than had previously been realized. Previously, biological oceanographers had focused on silicaceous diatoms as being the most important phytoplankton, accounting for 10-20 gigatons of carbon uptake each year. Chisholm's work showed that an even larger amount of carbon was cycled through these small algae, which may also play an important role in the global nitrogen cycle . In recent years, Chisholm has played a visible role in opposing the use of iron fertilization as a technological fix for anthropogenic climate change .Awards and honors Chisholm has been a member of the United States National Academy of Sciences since 2003 and a fellow of the American Academy of Arts and Sciences since 1992. In January 2010 , she was awarded the Alexander Agassiz Medal , for "pioneering studies of the dominant photosynthetic organisms in the sea and for integrating her results into a new understanding of the global ocean." She was a co-recipient in 2012 of the Ruth Patrick Award from the Association for the Sciences of Limnology and Oceanography . Chisholm received the National Medal of Science from President Barack Obama on February 1 , 2013. In 2013, she was awarded the Ramon Margalef Prize in Ecology , “for being one of the most productive, charismatic and active researchers on biology and marine ecology”. On May 24, 2018, she was awarded the Doctor of Science degree by Harvard University . In 2019 she received the Crafoord Prize in Biosciences, “for the discovery and pioneering studies of the most abundant photosynthesising organism on Earth, Prochlorococcus”. This prize is considered equivalent to the Nobel Prize . Chisholm was honored at the Crafoord Prize Symposium in Biosciences at which 6 internationally prominent scientists spoke : Alexandra Worden , Corina Brussaard , Ramunas Stepanauskas, Rachel Foster, Francis M . Martin and David Karl.Select works Chisholm, S.W. 2012. Unveiling Prochlorococcus: The Life and times of the ocean's smallest photosynthetic cell. 2012. In: Microbes and Evolution: The World That Darwin Never Saw. In: R. Kolter and S. Maloy . ASM Press. p. 165.
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