Ripiphoridae Ripiphoridae is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic parasitoids, an attribute that they share with the Meloidae . Members of the family differ in their choice of hosts , but most attack various species of bees or wasps, while some others attack cockroaches. Many species of Ripiphoridae have abbreviated elytra , and or antennae . Genera include Allocinops , Rhipistena and Sharpides . Species that attack bees typically lay their eggs on flowers. There the eggs hatch almost immediately into small planidial larvae and lie in wait for a visiting host. The planidium mounts the bee and rides it back to the hive . There it dismounts and seeks a cell occupied by a host larva. The planidium then enters the body of the host . It changes its skin and shape, then remains more or less dormant until the host larva pupates. It then emerges from the bee pupa and begins to feed. It eats the entire pupa, then pupates in its turn and completes its metamorphosis before emerging from the hive to mate and lay eggs. Fossil species in the genera Paleoripiphorus , Macrosiagon , Cretaceoripidius , Flabellotoma , Burmitoma , Plesiotoma , and Amberocula have been described from mid- to lower-Cretaceous amber from sites in France, Germany and Myanmar.Genera These 32 genera belong to the family Ripiphoridae: Alloclinops Broun, 1921 g Amberocula Batelka, Engel, & Prokop , 2018 g Ancholaemus Gerstaecker, 1855 g Blattivorus Chobaut, 1891 g Burmitoma Batelka, Engel, & Prokop, 2018 g Clinopalpus Batelka, 2009 g Clinops Gerstaecker, 1855 g Cretaceoripidius Falin & Engel, 2010 g Flabellotoma Batelka, Prokop, & Engel, 2016 g Geoscopus Gerstaecker, 1855 g Macrosiagon Hentz, 1830 i c g b Metoecus Dejean , 1834 g Micholaemus Viana, 1971 g Neopauroripidius Falin & Engel, 2014 g Neorrhipidius Viana, 1958 g Ohananomia Tôyama, 1986 g Paleoripiphorus Perrichot, Nel & Neraudeau, 2004 g Pauroripidius Kaupp & Nagel, 2001 g Pelecotoma Fischer, 1809 i c g b Pirhidius Besuchet, 1957 b Plesiotoma Batelka, Engel, & Prokop, 2018 g Ptilophorus Dejean, 1834 i c g b Quasipirhidius Zaragoza, 1992 g Quasirhipidius Zaragoza, 1992 g Rhipidocyrtus Falin & Engel, 2014 g Rhipistena Sharp, 1878 g Rhizostylops Silvestri , 1906 g Ripidius Thunberg , 1806 i c g Ripiphorus Bosc, 1791 i c g b Scotoscopus Reitter, 1884 g Trigonodera Dejean, 1834 i c g b Zapotecotoma Engel, Falin, & Batelka, 2019 i c g b Data sources: i = ITIS, c = Catalogue of Life , g = GBIF , b = Bugguide.net
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