Pennales


The order Pennales is a traditional subdivision of the heterokont algae known as diatoms. The order is named for the shape of the cell walls of pennate diatoms, which are elongated in valve view. The valves may be linear or oval in shape, and usually bear bilaterally symmetrical ornamental patterns. These patterns are composed of a series of transverse lines that can appear as rows of dots when viewed with an optical microscope. Some pennate diatoms also exhibit a fissure along their longitudinal axis. This is known as a raphe, and is involved in gliding movements made by diatom cells; motile diatoms always possess a raphe.
In terms of cell cycle, vegetative cells are diploid and undergo mitosis during normal cell division. Periodically, meiosis produces morphologically identical haploid gametes, which fuse to produce a zygote that develops into an auxospore.
In some taxonomic schemes, the pennate diatoms are divided into two groups: pennate diatoms without a raphe, known as araphids, and pennate diatoms with a raphe, known as raphids.