Issidae Issidae is a family of planthoppers described by Spinola in 1839, belonging to the order Hemiptera , suborder Auchenorrhyncha superfamily Fulgoroidea .Distribution Species of this family are present throughout the Northern Hemisphere .Description Issidae are small insects generally with a stocky body, as the wings mainly develop in width. Basic body coloration is not striking, usually shows brownish colors. The head has two ocelli . The forewings have strong pronounced ribs. They wrap the abdomen when the insect is at rest . The family originally included approximately 1000 species with 215 genera, but the systematics of Issidae remains uncertain, with many of the subfamilies having been recently removed to separate families, including Caliscelidae . Nogodinidae , and Tropiduchidae . In 2013, scientists described a biologically unique set of mechanical gears in an Issus nymph , though identical structures are known in most planthoppers, and were known for decades before the function of the gears was discovered Taxonomy The family Issidae was once large and included many groups which are now treated in other families or as families themselves. These groups include the Caliscelidae, Nogodinidae, and Tropiduchidae. Around 2003, there was a view in favour of a single subfamily Issinae, but the current consensus is placement in four. The Catalogue of Life lists genera in five tribes Issini, Parahiraciini , Hemisphaeriini and Thioniini . The tribe Colpopterini have now been placed in family Nogodinidae Melichar , 1898 having been raised to a subfamily.Subfamilies, Tribes and selected Genera Fulgoromorpha Lists On the Web lists four subfamilies:Authority: Melichar, 1906 Hemisphaeriini Melichar, 1906 * Gergithus * Hemisphaerius Kodaianellini Wang, Zhang & Bourgoin, 2016 Parahiraciini Cheng & Yang, 1991 Sarimini Wang, Zhang & Bourgoin, 2016 Hemisphaeriinae incertae sedis Hysteropterinae Authority: Melichar, 1906; there are currently 61 genera including: Agalmatium Emeljanov, 1971 Falcidius Stål, 1866 Hysteropterum Amyot & Audinet-Serville, 1843 - type genusIssinae The monotypic subfamily contains tribe Issini Spinola, 1839 and has a widespread distribution in Europe, the Middle East and temperate Asia. There are currently four genera: †Issites Haupt, 1956 Issus Brullé, 1832 – type genus Latissus Dlabola, 1974 Sinonissus Wang, Shi & Bourgoin, 2018Thioniinae Authority: Melichar, 1906 There are currently 3 tribes: Cordelini Gnezdilov, 2019 * Cordela Gnezdilov, 2019 Guianaphrynini Gnezdilov, 2018 * Guianaphryna Gnezdilov, 2018 Thioniini Melichar, 1906 * Oronoqua Fennah, 1947 * Cheiloceps Uhler, 1895 * Thionia Stal, 1859 * Waorania Gnezdilov & Bartlett, 2018 Twenty genera are currently placed here: Abolloptera Gnezdilov & O'Brien , 2006 Argepara Gnezdilov & O'Brien, 2008 Aztecus Gnezdilov & O'Brien, 2008 Balduza Gnezdilov & O'Brien, 2006 †Breukoscelis Szwedo, 2019 Bumaya Gnezdilov & O'Brien, 2008 Carydiopterum Gnezdilov, 2017 Delongana Caldwell, 1945 Exortus Gnezdilov, 2004 Gilda Walker , 1870 Incasa Gnezdilov & O'Brien, 2008 Kathleenum Gnezdilov, 2004 Paralixes Caldwell, 1945 Sarnus Stål, 1866 Stilbometopius Gnezdilov & O'Brien, 2006 Traxanellus Caldwell, 1945 Traxus Metcalf, 1923 Tylanira Ball, 1936 Ulixes Stål, 1861 Ulixoides Haupt, 1918
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