Prehistoric earthworks in the parish include the long barrow known as Tow Barrow, on Wexcombe Down, south of Wexcombe. Neolithic pottery was found in 1914 when the site was partially excavated by Crawford and Hooton. Marten is one of several possible sites for the murder of Cynewulf of Wessex in 786, and one of several suggested sites for the Battle of Marton in 871, in which Æthelred of Wessex suffered a defeat by the Viking army. A Roman road between Cirencester and Winchester passes Wilton and Marten. The area was part of the ancient parish of Great Bedwyn, formed from a large estate called Bedwyn which was recorded in 968. The 1086 Domesday Book recorded 16 households at Grafton and six at Marten. Earthworks at Marten, including evidence of a moat, are listed as a deserted medieval village. The Manor Farmhouses at West Grafton and Wilton are from the 17th century; Wexcombe Manor is from the 18th. The southern half of Great Bedwyn parish became a separate ecclesiastical district in 1844, when the church was built at East Grafton; the civil parish of Grafton was created in 1895. Having no large settlements, the population of the parish changed little in the 20th century. The manor of Wolfhall was transferred to Burbage parish in 1988.
The Church of England parish church of St Nicholas at East Grafton was built in 1842-44, to designs by Benjamin Ferrey; before then the parish church was at Great Bedwyn. In 1986 the church was designated as Grade II* listed. East Grafton, Marten and Wilton each had a chapel of ease in the 14th and 15th centuries, and possibly earlier. Wexcombe had a chapel of ease by 1879, where services were conducted by the vicar of Tidcombe from 1899; it closed in the 1920s. In Tidcombe churchyard are 19th-century tombs of members of the Hawkins family of Wexcombe. A Wesleyan Methodist chapel was built at Wilton in 1811, extended with a schoolroom in 1843 and improved in the 1860s. It closed in 1994 and is in residential use. Primitive Methodist chapels were built at West Grafton by 1874 and at Wexcombe in the 1880s. By the 1960s, both were in residential use.
East Grafton has village hall, the Coronation Hall, built in 1937 and renovated in 2009. There is no school in the parish. A National School was opened at East Grafton in 1846 and improved in the 20th century; it closed in 2011 owing to falling pupil numbers. Wilton has a pub, The Swan. On high ground above Wilton is a working 19th-century flour mill, Wilton Windmill. Crofton Pumping Station, which supplies the Kennet and Avon Canal and has a 200-year-old beam engine, is just beyond the northern boundary of the parish. Its reservoir, Wilton Water, is a nature reserve.
