According to his service record, which has been placed online by The National Archives, Prowse was born on 28 August 1896, however the Commonwealth War Graves Commission gives his age at death as 32, which would place his birth in 1886. Recent research in South Wales has unearthed a birth certificate that shows he was actually born on 29 August 1886 in Brynsion Terrace, Gilfach Goch, Llantrisant and a Blue Plaque was placed on the house by the local council, Rhondda Cynon Taff on 21 October 2006. He enlisted in the Royal Navy Volunteer Reserve for the Royal Naval Division, on 26 February 1915. The details recorded in his service record show he was living in the Landore area of Swansea, Wales with his wife Sarah; he had been working as a collier, was tall, had grey eyes, brown hair and a "fair" complexion. At some time he relocated to Camerton in Somerset, probably to work in the coal mines there. In the churchyard at Camerton there is a war memorial with his name shown. Prowse was initially ratedOrdinary Seaman, on 5 May 1915 he was promoted to Able Seaman. After completing training at Blandford, in September 1915 he was posted to the Drake Battalion, which was then engaged in the Gallipoli Campaign. It is not clear how much active service Prowse saw at this time as he spent significant periods hospitalised with first jaundice and then gastroenteritis, finally rejoining his battalion in Egypt on 9 January 1916. The division remained engaged in the Gallipoli Campaign until May 1916 when it was transferred to France. Prowse arrived at Marseilles on 7 June 1916, on 20 June he was promoted Petty Officer. In November 1916 the Division was employed in the Battle of the Ancre, the final attempt to resolve the Battle of the Somme. Prowse received a gunshot wound in his left thigh on 13 November and after initial treatment in France was admitted to a hospital in Epsom on 17 November. Having been discharged from hospital, he went back to the base at Blandford on 9 January 1917, and eventually returned to his battalion in France on 28 March 1917. In August 1918 Prowse was recommended for the award of the DCM. The citation, published posthumously in 1919 stated Prowse was about 32 years old, and a Chief Petty Officer in the Royal Naval Volunteer Reserve, during the First World War when the following deed took place for which he was awarded the VC. On 2 September 1918 at Pronville, France, Chief Petty Officer Prowse led a small party of men against an enemy strong-point, capturing it, together with 23 prisoners and 5 machine-guns. On three other occasions he displayed great heroism in dealing with difficult and dangerous situations, and at one time he dashed forward and attacked and captured two machine-gun posts, killing six of the enemy and taking 13 prisoners and two machine-guns. He was the only survivor of this gallant party, but his action enabled the battalion to push forward in comparative safety.
Sarah Prowse was presented with her husband's Victoria Cross by H.M. King George V at Buckingham Palace on 17 July 1919, after all the live recipients. She led the list of posthumous awards and was presented with the last VC with the blue naval ribbon. Prowse's Victoria Cross is in the Lord Ashcroft VC Collection at the Imperial War Museum.