Forfarella


Forfarella is a genus of chasmataspidid, a group of extinct aquatic arthropods.

Description

Like the other chasmataspidids, F. mitchelli was a small arthropod, with the size of its only known specimen estimated at only.
Its carapace was wide and rectangular. It was broader than long; wide and long. Although the details on its surface are little preserved, a tubercle is distinguished in the left anterior corner of the carapace if it is placed under plane polarized light. This tubercle most likely represents a lateral eye. The appendages are only known from two fragments on both left and right sides, probably belonging to the sixth pair of them. The left one had a length of and a width of, while the right one was long and wide.
The opisthosoma, made up of 13 segments, had a total length of. The preabdomen was subtrapezoidal and narrowed posteriorly. Vague impressions distinguish the second, third and fourth tergites, but not the first one. The postabdomen also tapered posteriorly and was longer, with its 9 tergites distinguishable in the fossil and with a length of approximately each. The telson is unknown, but it was probably short as in other contemporary chasmataspidids. The specimen had two cracks all over the prosoma and preabdomen, as well as dark patches probably corresponding to the ornamentation.

History of resarch

Forfarella is known from one single specimen, BMNH In 60023. It was found in the Old Red Sandstone near Arbroath, Scotland, in the United Kingdom. According to the label associated with the fossil, it was purchased in 1893 from the collection of a person surnamed Mitchell, who, along with other colleagues, was one of the fossil collectors in the area who did not usually specify the exact type locality of his material. This rendered its tracking problematic, but comparisons with other fossils from Mitchell's collection that had more details about its origins and the fossil's label suggest the Kelly Den south of Arbirlot, although this cannot be confirmed. Following the purchase, the fossil of Forfarella was not instantly registered in the Natural History Museum of London like other fossils of Mitchell's collection.
In 1962, the British paleontologist Charles D. Waterston borrowed the still unregistered In 60023, as well as four other fossils of juvenile eurypterids, to describe them. He noted the chasmataspidid nature of In 60023, and coined the name Forfarella mitchelli. However, Waterston would never publish his description, returning the fossil to the museum three years later, where it would be labeled with the name given by the paleontologist. Thus, Forfarella mitchelli remained as a nomen manuscriptum. But despite the turbulent history of the fossil, the paleontologists Jason A. Dunlop, Lyall I. Anderson and Simon J. Braddy formally proposed and described Forfarella mitchelli for the first time in 1999 as a new genus of chasmataspidid. The name suggested by Waterston was kept to avoid future confusion. Anderson also studied a troublesome specimen allegedly at the National Museum of Scotland, which could represent another specimen of Forfarella. However, upon examination, it was discovered that it was not a chasmataspidid, but a poorly preserved undetermined arthropod fragment.

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