Flower chafer
Flower chafers are a group of scarab beetles, comprising the subfamily Cetoniinae. Many species are diurnal and visit flowers for pollen and nectar, or to browse on the petals. Some species also feed on fruit. The group is also called fruit and flower chafers, flower beetles and flower scarabs. There are around 4,000 species, many of them still undescribed.
Overview
By morphological characters, the adults can be separated from the other scarabs by the combination of the following characters: epipleuron easily recognizable, border lateral of elytra sinuate and antennal insertion visible from above. Twelve tribes are presently recognized: Cetoniini, Cremastocheilini, Diplognathini, Goliathini, Gymnetini, Phaedimini, Schizorhinini, Stenotarsiini, Taenioderini, Trichiini, Valgini, and Xiphoscelidini. The tribe Gymnetini is the biggest of the American tribes, and Goliathini contains the largest species, and is mainly found in the rainforest regions of Africa.Biology
Many species in the tribe Cremastocheilini are known to be predaceous, feeding on hymenopteran larvae or soft-bodied nymphs of Auchenorrhyncha. Spilophorus spp. have been noted feeding on the nesting material and excrement of South African passerine birds, while Spilophorus maculatus has been recorded feeding on Oxyrhachis sp. nymphs and Hoplostomus fuligineus is known to feed on the brood of honey bees in South Africa and the pupae of the wasp Belonogaster petiolata. Campsiura javanica feeds on the larvae of Ropalidia montana in southern India. Cremastocheilus stathamae feeds on ants of the genus Myrmecocystus.Systematics and taxonomy
The tribes of subfamily Cetoniinae, with some notable genera also listed, are:Tribus [Cetoniini]
Subtribus Cetoniina- Aethiessa Burmeister, 1842
- Anatona Burmeister, 1842
- Anelaphinis Kolbe, 1912
- Aphelinis Antoine, 1987
- Atrichelaphinis Kraatz, 1898
- Atrichiana Distant, 1911
- Badizoblax Thomson, 1877
- Callophylla Moser, 1916
- Centrantyx Fairmaire, 1884
- Cetonia Fabricius, 1775
- Chiloloba Burmeister, 1842
- Cosmesthes Kraatz, 1880
- Cosmiophaena Kraatz, 1899
- Dischista Burmeister, 1842
- Dolichostethus Kolbe, 1912
- Elaphinis Burmeister, 1842
- Enoplotarsus Lucas, 1859
- Erlangeria Preiss, 1902
- Euglypta Mohnike, 1873
- Gametis Burmeister, 1842
- Gametoides Antoine, 2006
- Glycosia Schoch, 1896
- Glycyphana Burmeister, 1842
- Gymnophana Arrow, 1910
- Hemiprotaetia Mikšič, 1963
- Heteralleucosma Antoine, 1989
- Heterocnemis Albers, 1852
- Heterotephraea Antoine, 2002
- Latescutella Ruter, 1972
- Lawangia Schenkling, 1921
- Lorkovitschia Mikšič, 1968
- Marmylida Thomson, 1880
- Micrelaphinis Schoch, 1896
- Miksicus Özdikmen & Turgat, 2009
- Mireia Ruter, 1953
- Niphobleta Kraatz, 1880
- Pachnoda Burmeister, 1842
- Paleopragma Thomson, 1880
- Paralleucosma Antoine, 1989
- Paranelaphinis Antoine, 1988
- Paraprotaetia Moser, 1907
- Pararhabdotis Kraatz, 1899
- Parastraella Antoine, 2006
- Parelaphinis Holm & Marais, 1989
- Phaneresthes Kraatz, 1894
- Phonotaenia Kraatz, 1880
- Phoxomeloides Schoch, 1898
- Podopholis Moser, 1915
- Podopogonus Moser, 1917
- Polybaphes Kirby, 1827
- Polystalactica Kraatz, 1882
- Protaetia Burmeister, 1842
- *Liocola Thomson, 1859
- *Potosia Mulsant & Rey, 1871
- Protaetiomorpha Mikšič, 1968
- Psacadoptera Kraatz, 1882
- Pseudoprotaetia Kraatz, 1882
- Pseudotephraea Kraatz, 1882
- Pseudourbania Mikšič, 1965
- Reineria Mikšič, 1968
- Rhabdotis Burmeister, 1842
- Rhabdotops Krikken, 1981
- Rhyxiphloea Burmeister, 1842
- Simorrhina Kraatz, 1886
- Somalibia Lansberge, 1882
- Stalagmosoma Burmeister, 1842
- Sternoplus Wallace, 1868
- Systellorhina Kraatz, 1895
- Tephraea Burmeister, 1842
- Thyreogonia Reitter, 1898
- Trichocelis Moser, 1908
- Trichocephala Moser, 1916
- Trichostetha Burmeister, 1842
- Tropinota Mulsant, 1842
- Urbania Mikšič, 1963
- Xeloma Kraatz, 1881
- Xiphosceloides Holm, 1992
- Euphoria Burmeister, 1842
- Chlorixanthe Bates, 1889
- Acrothyrea Kraatz, 1882
- Alleucosma Schenkling, 1921
- Analleucosma Antoine, 1989
- Cyclophorellus Krikken, 1982
- Cyrtothyrea Kolbe, 1895
- Discopeltis Burmeister, 1842
- Grammopyga Kraatz, 1895
- Homothyrea Kolbe, 1895
- Leucocelis Burmeister, 1842
- Leucochilus Kraatz, 1896
- Lonchothyrea Kolbe, 1895
- Mausoleopsis Lansberge, 1882
- Mecaspidiellus Antoine, 1997
- Mecaspidius Bourgoin, 1921
- Megalleucosma Antoine, 1989
- Molynoptera Kraatz, 1897
- Molynopteroides Antoine, 1989
- Oxythyrea Mulsant, 1842
- Paleira Reiche, 1871
- Phoxomela Schaum, 1844
- Pseudalleucosma Antoine, 1989
- Pseudooxythyrea Baraud, 1985
- Trichothyrea Kolbe, 1895
- Chewia Legrand, 2004
- Pachnodoides Alexis & Delpont, 2002
Tribus [Cremastocheilini]
- Aspilus Schaum, 1848
- Protochilus Krikken, 1976
- Arielina Rossi, 1958
- Astoxenus Péringuey, 1907
- Basilewskynia Schein, 1957
- Coenochilus Schaum, 1841
- Ruterielina Rojkoff, 2010
- Centrochilus Krikken, 1976
- Clinterocera Motschulsky, 1857
- Cremastocheilus Knoch, 1801
- Cyclidiellus Krikken, 1976
- Cyclidinus Westwood, 1874
- Cyclidius MacLeay, 1838
- Genuchinus Westwood, 1874
- Lissomelas Bates, 1889
- Paracyclidius Howden, 1971
- Platysodes Westwood, 1873
- Psilocnemis Burmeister, 1842
- Cymophorus Kirby, 1827
- Myrmecochilus Wasmann, 1900
- Rhagopteryx Burmeister, 1842
- Genuchus Kirby, 1825
- Meurguesia Ruter, 1969
- Problerhinus Deyrolle, 1864
- Goliathopsis Janson, 1881
- Heterogenius Moser, 1911
- Chtonobius Burmeister, 1847
- Lissogenius Schaum, 1844
- Brachymacroma Kraatz, 1896
- Campsiura Hope, 1831
- Macromina Westwood, 1874
- Pseudopilinurgus Moser, 1918
- Nyassinus Westwood, 1879
- Anatonochilus Péringuey, 1907
- Laurentiana Ruter, 1952
- Oplostomus W.S. MacLeay, 1838
- Placodidus Péringuey, 1900
- Scaptobius Schaum, 1841
- Callynomes Mohnike, 1873
- Centrognathus Guérin-Méneville, 1840
- Parapilinurgus Arrow, 1910
- Pilinurgus Burmeister, 1842
- Spilophorus Schaum, 1848
- Telochilus Krikken, 1975
- Lecanoderus Kolbe, 1908
- Trichoplus Burmeister, 1842
- Pseudoscaptobius Krikken, 1976
- Trogodes Westwood, 1873
Tribus [Diplognathini]
- Anthracophora Burmeister, 1842
- Anthracophorides Moser, 1918
- Apocnosis J. Thomson, 1878
- Apocnosoides Antoine, 2001
- Charadronota Burmeister, 1842
- Conradtia Kolbe, 1892
- Diphrontis Gerstäcker, 1882
- Diplognatha Gory & Percheron, 1833
- Eriulis Burmeister, 1842
- Hadrodiplognatha Kraatz, 1898
- Heteropseudinca Valck Lucassen, 1933
- Metallopseudinca Valck Lucassen, 1933
- Niphetophora Kraatz, 1883
- Parapoecilophila Hauser, 1904
- Pilinopyga Kraatz, 1880
- Poecilophila Kraatz, 1893
- Poecilophilides Kraatz, 1898
- Porphyronota Burmeister, 1842
- Pseudinca Kraatz, 1880
- Ruteroides Alves, 1973
- Stethopseudinca Valck Lucassen, 1933
- Uloptera Burmeister, 1842
Tribus [Goliathini]
- Aphelorrhina Westwood, 1841
- Asthenorhella Westwood, 1874
- Asthenorrhina Westwood, 1843
- Bothrorrhina Burmeister, 1842
- Caelorrhina Hope, 1841
- Cheirolasia Westwood, 1843
- Chlorocala Kirby & Spence, 1828
- Cyphonocephalus Westwood, 1842
- Cyprolais Burmeister, 1842
- Dicheros Gory & Percheron, 1833
- Dicronorhina Hope, 1837
- Eudicella White, 1839
- Genyodonta Burmeister, 1842
- Gnathocera Kirby, 1825
- Heterorrhina Westwood, 1842
- Ischnoscelis Burmeister, 1842
- Mecynorhina Hope, 1837
- *Amaurodes Westwood, 1844
- *Megalorhina Westwood, 1847
- Mystroceros Burmeister, 1842
- Neoscelis Schoch, 1897
- Plaesiorrhina Burmeister, 1842
- Pseudotorynorrhina Mikšič, 1967
- Rhomborhina Hope 1837
- Stephanorrhina Burmeister, 1842
- Smicorhina Westwood, 1847
- Taurhina Burmeister, 1842
- *Neptunides J. Thomson, 1879
- *Rhamphorrhina Klug, 1855
- Tmesorrhina Westwood, 1842
- Torynorrhina Arrow, 1907
- Dicronocephalus Hope, 1837
- Platynocephalus Westwood, 1854
- Argyrophegges Kraatz, 1895
- Fornasinius Bertoloni, 1853
- Goliathus Lamarck, 1801
- Hegemus J. Thomson, 1881
- Hypselogenia Burmeister, 1840
- Ichnestoma Gory & Percheron
- Anisorrhina Westwood, 1842
- Aphanochroa Kolbe, 1893
- Bietia Fairmaire, 1898
- Brachymitra Kolbe, 1904
- Ceratorrhina Westwood, 1843
- Chondrorrhina Kraatz, 1880
- Chordodera Burmeister, 1842
- Compsocephalus White, 1845
- Cosmiomorpha Saunders, 1854
- Daedycorrhina Bates, 1888
- Dicellachilus Waterhouse, 1905
- Dyspilophora Kraatz, 1880
- Eutelesmus Waterhouse, 1880
- Gnorimimelus Kaatz, 1880
- Herculaisia Seilliere, 1910
- Inhambane Péringuey, 1907
- Jumnos Saunders, 1839
- Lophorrhina Westwood, 1842
- Melinesthes Kraatz, 1880
- Narycius Dupont, 1835
- Neophaedimus Schoch, 1894
- Neoscelis Schoch, 1897
- Pedinorrhina Kraatz, 1880
- Priscorrhina Krikken, 1984
- Ptychodesthes Kraatz, 1883
- Raceloma J. Thomson, 1877
- Scythropesthes Kraatz, 1880
- Smaragdesthes Kraatz, 1880
- Spelaiorrhina Lansberge, 1886
- Stephanocrates Kolbe, 1892
- Taeniesthes Kraatz, 1880
- Trigonophorus'' Hope, 1831
Tribus [Gymnetini]
- Blaesia Burmeister, 1842
- Halffterinetis Morón and Nogueira, 2007
- Aemilius LeMoult, 1939
- Allorhina Burmeister, 1842
- Amazula Kraatz, 1882
- Amithao J. Thomson, 1878
- Argyripa J. Thomson, 1878
- Astrocara Schürhoff, 1937
- Badelina J. Thomson, 1880
- Balsameda J. Thomson, 1880
- Chiriquiba Bates, 1889
- Cineretis Schürhoff, 1937
- Clinteria Burmeister, 1842
- Clinteroides Schoch, 1898
- Cotinis Burmeister, 1842
- Desicasta J. Thomson, 1878
- Guatemalica Poll, 1886
- Gymnetina Casey, 1915
- Gymnetis MacLeay, 1819
- Gymnetosoma Martínez, 1949
- Hadrosticta Kraatz, 1892
- Heterocotinis Martínez, 1949
- Hoplopyga J. Thomson, 1880
- Hoplopygothrix Schürhoff, 1933
- Jansonella Blackwelder, 1944
- Macrocranius Schürhoff, 1935
- Marmarina Kirby, 1827
- Neocorvicoana Ratcliffe & Micó, 2001
- Stethodesma Bainbridge, 1840
- Tiarocera'' Burmeister, 1842
Tribus [Phaedimini]
- Phaedimus Waterhouse, 1841
- Hemiphaedimus Mikšič, 1972
- Philistina W.S. MacLeay, 1838
- Theodosia J. Thomson, 1880
Tribus [Schizorhinini]
- Agestrata Eschscholtz, 1829
- Ischiopsopha Gestro, 1874
- Lomaptera Gory & Percheron, 1833
- Lomapteroides Schoch, 1898
- Megaphonia Schürhoff, 1933
- Morokia Janson, 1905
- Mycterophallus Van de Poll, 1886
- Aphanesthes Kraatz, 1880
- Bislardiana Antoine, 2003
- Chalcopharis Heller, 1901
- Chlorobapta Kraatz, 1880
- Chondropyga Kraatz, 1880
- Clithria Burmeister, 1842
- Diaphonia Newman, 1840
- Dichrosoma Kraatz, 1885
- Dilochrosis Thomson, 1878
- Eupoecila Burmeister, 1842
- Grandaustralis Hutchinson & Moeseneder, 2013
- Hemichnoodes Kraatz, 1880
- Hemipharis Burmeister, 1842
- Lenosoma Kraatz, 1880
- Lyraphora Kraatz, 1880
- Macrotina Strand, 1934
- Metallesthes Kraatz, 1880
- Micropoecila Kraatz, 1880
- Neoclithria Van de Poll, 1886
- Neorrhina J. Thomson, 1878
- Octocollis Moeseneder & Hutchinson, 2012
- Phyllopodium Schoch, 1895
- Poecilopharis Kraatz, 1880
- Pseudoclithria Van de Poll, 1886
- Schizorhina Kirby, 1825
- Schochidia Berg, 1898
- Stenopisthes Moser, 1913
- Tapinoschema Thomson, 1880
- Trichaulax Kraatz, 1880
Tribus [Stenotarsiini]
- Anochilia Burmeister, 1842
- Epistalagma Fairmaire, 1880
- Chromoptilia Westwood, 1842
- Descarpentriesia Ruter, 1964
- Bricoptis Burmeister, 1842
- Coptomia Burmeister, 1842
- Coptomiopsis Pouillaude, 1919
- Euchilia Burmeister, 1842
- Euryomia Burmeister, 1842
- Heterocranus Bourgoin, 1919
- Hiberasta Fairmaire, 1901
- Hyphelithia Kraatz, 1880
- Liostraca Burmeister, 1842
- Micreuchilia Pouillaude, 1917
- Micropeltus Blanchard, 1842
- Pareuchilia Kraatz, 1880
- Pseudeuryomia Kraatz, 1894
- Pygora Burmeister, 1842
- Pyrrhopoda Kraatz, 1880
- Vieuella Ruter, 1964
- Doryscelis Burmeister, 1842
- Epixanthis Burmeister, 1842
- Hemiaspidius Krikken, 1982
- Parepixanthis Kraatz, 1893
- Pseudepixanthis Kraatz, 1880
- Rhynchocephala Fairmaire, 1883
- Euchroea Burmeister, 1842
- Heterophana Burmeister, 1842
- Oxypelta Pouillaude, 1920
- Pogonotarsus Burmeister, 1842
- Zebinus Fairmaire, 1894
- Heterosoma Schaum, 1845
- Plochilia Fairmaire, 1896
- Bonoraella Ruter, 1978
- Celidota Burmeister, 1842
- Cyriodera Burmeister, 1842
- Dirrhina Burmeister, 1842
- Hemilia Kraatz, 1880
- Lucassenia Olsoufieff, 1940
- Moriaphila Kraatz, 1880
- Pantolia Burmeister, 1842
- Tetraodorhina Blanchard, 1842
- Parachilia Burmeister, 1842
- Callipechis Burmeister, 1842
- Ischnotarsia Kraatz, 1880
- Rhadinotaenia Kraatz, 1900
- Stenotarsia Burmeister, 1842
- Vadonidella Ruter, 1973
Tribus [Taenioderini]
- Anocoela Moser, 1914
- Bacchusia Mikšic, 1976
- Bombodes Westwood, 1848
- Carneluttia Mikšic, 1976
- Chalcothea Burmeister, 1842
- Chalcotheomima Mikšic, 1970
- Clerota Burmeister, 1842
- Coilodera Hope, 1831
- Costinota Schürhoff, 1933
- Eumacronota Mikšic, 1976
- Euremina Wallace, 1867
- Euselates Thomson, 1880
- Glyptothea Bates, 1889
- Glyptotheomima Mikšic, 1976
- Gnorimidia van Lansberge, 1887
- Hemichalcothea Mikšic, 1970
- Ixorida Thomson, 1880
- *Mecinonota Kraatz, 1892
- *Oncosterna Thomson, 1880
- *Pseudomecinonota Miksic, 1972
- Macronotops Krikken, 1977
- Meroloba Thomson, 1880
- Microchalcothea Moser, 1910
- Penthima Kraatz, 1892
- Plectrone Wallace, 1867
- Pleuronota Kraatz, 1892
- Pseudochalcothea Ritsema, 1882
- Taeniodera Burmeister, 1842
- Xenoloba Bates, 1889
Tribus [Trichiini]
Subtribus Cryptodontina- Coelocorynus Kolbe, 1895
- Cryptodontes Burmeister, 1847
- Golinca Thomson, 1878
- Inca Lepeletier & Serville, 1828
- Pantodinus Burmeister, 1847
- Osmoderma Lepeletier & Serville, 1825
- Platygeniops Krikken, 1978
- Platygenia MacLeay, 1819
- Agnorimus Miyake et al., 1991
- Apeltastes Howden, 1968
- Brachagenius Kraatz, 1890
- Calometopidius Bourgoin, 1917
- Calometopus Blanchard, 1850
- Campulipus Kirby, 1827
- Chaetodermina Heller, 1921
- Clastocnemis Burmeister, 1840
- Coelocratus Burmeister, 1841
- Corynotrichius Kolbe, 1891
- Dialithus Parry, 1849
- Diploa Kolbe, 1892
- Diploeida Péringuey, 1907
- Elpidus Péringuey, 1907
- Endoxazus Kolbe, 1892
- Epitrichius Tagawa, 1941
- Eriopeltastes Burmeister, 1840
- Giesbertiolus Howden, 1988
- Glaphyronyx Moser, 1924
- Gnorimella Casey, 1915
- Gnorimus Lepeletier & Serville, 1825
- Incala J. Thomson, 1858
- Incalidia Janson, 1907
- Liotrichius Kolbe, 1892
- Myodermides Ruter, 1964
- Myodermum Burmeister, 1840
- Paragnorimus Becker, 1910
- Paratrichius Janson, 1881
- Peltotrichius Howden, 1968
- Pileotrichius Bourgoin, 1921
- Polyplastus Janson, 1880
- Stegopterus Burmeister, 1840
- Stripsipher Gory & Percheron, 1833
- Trichiomorphus Bourgoin, 1919
- Trichiotinus Casey, 1915
- Trichius Fabricius, 1787
- Trigonopeltastes Burmeister, 1840
- Xiphoscelidus Péringuey, 1907
Tribus [Valgini]
- Ischnovalgus Kolbe, 1897
- Microvalgus Kraatz, 1883
- Stenovalgus Kolbe, 1892
- Acanthovalgus Kraatz, 1895
- Bivalgus Paulian, 1961
- Chaetovalgus Moser, 1914
- Charitovalgus Kolbe, 1904
- Chromovalgus Kolbe, 1897
- Comythovalgus Kolbe, 1897
- Cosmovalgus Kolbe, 1897
- Dasyvalgoides Endrödi, 1952
- Dasyvalgus Kolbe, 1904
- Euryvalgus Moser, 1908
- Excisivalgus Endrödi, 1952
- Heterovalgus Krikken, 1978
- Homovalgus Kolbe, 1897
- Hoplitovalgus Kolbe, 1904
- Hybovalgus Kolbe, 1904
- Idiovalgus Arrow, 1910
- Lepivalgus Moser, 1914
- Lobovalgus Kolbe, 1897
- Mimovalgus Arrow, 1944
- Oedipovalgus Kolbe, 1897
- Oreoderus Burmeister, 1842
- Oreovalgus Kolbe, 1904
- Podovalgus Arrow, 1910
- Pygovalgus Kolbe, 1884
- Sphinctovalgus Kolbe, 1904
- Tibiovalgus Kolbe, 1904
- Valgoides Fairmaire, 1899
- Valgus Scriba, 1790
- Xenoreoderus Arrow, 1910
- Yanovalgus Nomura, 1952
Tribus [Xiphoscelidini]
- Aporecolpa van Lansberge, 1886
- Callophylla Moser, 1916
- Heteroclita Burmeister, 1842
- Ischnostomiella Krikken, 1978
- Lansbergia Ritsema, 1888
- Meridioclita Krikken, 1982
- Myodermidius Bourgoin, 1920
- Oroclita Krikken, 1982
- Phonopleurus Moser, 1919
- Plochiliana Ruter, 1978
- Protoclita Krikken, 1978
- Rhinocoeta Burmeister, 1842
- *Haematonotus Kraatz, 1880
- Scheinia Ruter, 1958
- Xiphoscelis Burmeister, 1842