Curly-tailed lizard
The curly-tailed lizards belong to the family Leiocephalidae. One of the defining features of these lizards is that their tail often curls over. They were previously regarded as members of subfamily Leiocephalinae within the family Tropiduridae. There are presently 29 known species, all in the genus Leiocephalus.
Distribution
The curly-tailed lizards are native to the West Indies, with the extant species in the Bahamas, Turks and Caicos, Cayman Islands, Cuba, Hispaniola and nearby small islands. Additionally, Leiocephalus carinatus and Leiocephalus schreibersii have been introduced to Florida.General anatomy
The curly-tailed lizards vary in size depending on species, but typically are approximately in snout-to-vent length. These lizards have no femoral pores, pterygoid teeth, or palatine teeth. Additionally, these lizards are observed to have overlapping scales.Behavior
The curly-tailed lizards mostly forage on arthropods like insects, but also commonly take flowers and fruits. Large individuals will eat small vertebrates, including anoles.As suggested by their name, most species of this family often lift their tail and curl it. This is done both when a potential predator is present and when not present, although in some curly-tailed lizard species it increases when one is present. It shows the fitness of the lizard to a would-be predator and—in the case of an attack—draws attention to the tail, which increases the lizard's chance of escaping. Although it has been suggested that it also functions as a territorial display, studies have been unable to find support for this, as the tail curling does not change when another member of the same species is present.
Conservation status
The conservation status of the species in this family varies greatly. Several species, for example Leiocephalus carinatus, are common and widespread. Others are rare and highly threatened, especially those restricted to a single small island or a single location on a larger island, like the critically endangered Leiocephalus altavelensis from Alto Velo Island and critically endangered Leiocephalus onaneyi from Guantánamo Province in Cuba. Primary threats to their survival are habitat loss and introduced predators. Several species are already extinct. Some of these are only known from fossil or subfossil remains and became extinct in the Pleistocene or pre-Columbian era, but others survived into recent history. Three species, Leiocephalus endomychus, Leiocephalus pratensis and Leiocephalus rhutidira, have not been seen since the 1960s and 1970s and are recognized as critically endangered, possibly extinct, by the IUCN. They are among the "most wanted" EDGE species.Newly discovered species
Lizards of this family are diurnal and mostly inhabit fairly open habitats in a generally well-studied part of the world. Consequently, the majority of the species and subspecies already were scientifically described several decades ago. In 2016, the first new curly-tailed lizard since the early 1980s was described. The species was found in the coastal dunes of Bahía de las Calderas in the southwestern Dominican Republic. This species differs from the rest within Leiocephalidae in that its bony parietal table is U-shaped versus V-shaped, the males have 3-4 enlarged post-postcloacal scales versus 2, and there are specific sexual dimorphism trails.Species and subspecies
The following species and subspecies, listed alphabetically by scientific name, are recognized as being valid by the Reptile Database.- Leiocephalus barahonensis – orange-bellied curly-tailed lizard
- *Leiocephalus barahonensis altavelensis – Alto Velo curly-tailed lizard
- *Leiocephalus barahoensis aureus
- *Leiocephalus barahonensis barahonensis
- *Leiocephalus barahoensis beatanus
- *Leiocephalus barahoensis oxygaster
- Leiocephalus carinatus – Cuban curly-tailed lizard, northern curly-tailed lizard
- *Leiocephalus carinatus aquarius
- *Leiocephalus carinatus armouri – Little Bahama curly-tailed lizard
- *Leiocephalus carinatus carinatus
- *Leiocephalus carinatus cayensis
- *Leiocephalus carinatus coryi – saw-scaled curly-tailed lizard
- *Leiocephalus carinatus granti – Cayman Brac curly-tailed lizard
- *Leiocephalus carinatus hodsoni
- *Leiocephalus carinatus labrossytus
- *Leiocephalus carinatus microcyon
- *Leiocephalus carinatus mogotensis
- *Leiocephalus carinatus virescens
- *Leiocephalus carinatus zayasi
- Leiocephalus cubensis – Cuban brown curly-tailed lizard
- *Leiocephalus cubensis cubensis
- *Leiocephalus cubensis gigas
- *Leiocephalus cubensis minor
- *Leiocephalus cubensis pambasileus
- *Leiocephalus cubensis paraphrus
- † Leiocephalus cuneus – Antiguan curly-tailed lizard
- Leiocephalus endomychus – Hinche curly-tailed lizard
- † Leiocephalus eremitus – Navassa curly-tailed lizard
- † Leiocephalus etheridgei – Etheridge's curly-tailed lizard
- Leiocephalus greenwayi – East Plana curly-tailed lizard
- † Leiocephalus herminieri – Martinique curly-tailed lizard
- Leiocephalus inaguae – Inagua curly-tailed lizard
- † Leiocephalus jamaicensis – Jamaican curly-tailed lizard
- Leiocephalus loxogrammus – San Salvador curly-tailed lizard
- *Leiocephalus loxogrammus loxogrammus
- *Leiocephalus loxogrammus parnelli
- Leiocephalus lunatus – Santo Domingo curly-tailed lizard
- *Leiocephalus lunatus arenicolor
- *Leiocephalus lunatus lewisi
- *Leiocephalus lunatus louisae
- *Leiocephalus lunatus lunatus
- *Leiocephalus lunatus melaenoscelis
- *Leiocephalus lunatus thomasi
- Leiocephalus macropus – Cuban side-blotched curly-tailed lizard
- *Leiocephalus macropus aegialus
- *Leiocephalus macropus asbolomus
- *Leiocephalus macropus felinoi
- *Leiocephalus macropus hoplites
- *Leiocephalus macropus hyacinthurus
- *Leiocephalus macropus immaculatus
- *Leiocephalus macropus koopmani
- *Leiocephalus macropus lenticulatus
- *Leiocephalus macropus macropus
- *Leiocephalus macropus phylax
- *Leiocephalus macropus torrei
- Leiocephalus melanochlorus – Tiburon curly-tailed lizard
- *Leiocephalus melanochlorus hypsistus
- *Leiocephalus melanochlorus melanochlorus
- Leiocephalus onaneyi – Guantanamo striped curly-tailed lizard
- † Leiocephalus partitus
- Leiocephalus personatus – Hispaniolan masked curly-tailed lizard
- Leiocephalus pratensis – Haitian striped curly-tailed lizard
- *Leiocephalus pratensis chimarus
- *Leiocephalus pratensis pratensis
- Leiocephalus psammodromus – Turks and Caicos curly-tailed lizard
- *Leiocephalus psammodromus aphretor
- *Leiocephalus psammodromus apocrinus
- *Leiocephalus psammodromus cacodoxus
- *Leiocephalus psammodromus hyphantus
- *Leiocephalus psammodromus mounax
- *Leiocephalus psammodromus psammodromus
- Leiocephalus punctatus – Crooked Acklins curly-tailed lizard
- Leiocephalus raviceps – pallid curly-tailed lizard
- *Leiocephalus raviceps delavarai
- *Leiocephalus raviceps jaumei
- *Leiocephalus raviceps kilinikowski
- *Leiocephalus raviceps raviceps
- *Leiocephalus raviceps uzzelli
- Leiocephalus rhutidira – black-throated curly-tailed lizard
- Leiocephalus schreibersii – red-sided curly-tailed lizard
- *Leiocephalus schreibersii nesomorus
- *Leiocephalus schreibersii schreibersii
- Leiocephalus semilineatus – pale-bellied curly-tailed lizard
- Leiocephalus sixtoi
- Leiocephalus stictigaster – Cuban striped curly-tailed lizard
- *Leiocephalus stictigaster astictus
- *Leiocephalus stictigaster celeustes
- *Leiocephalus stictigaster exotheotus
- *Leiocephalus stictigaster gibarensis
- *Leiocephalus stictigaster lipomator
- *Leiocephalus stictigaster lucianus
- *Leiocephalus stictigaster naranjoi
- *Leiocephalus stictigaster ophiplacodes
- *Leiocephalus stictigaster parasphex
- *Leiocephalus stictigaster septentrionalis
- *Leiocephalus stictigaster sierrae
- *Leiocephalus stictigaster stictigaster
- Leiocephalus varius – Grand Cayman curly-tailed lizard
- Leiocephalus vinculum – Cochran's curlytail lizard