Villiers was the third son of Sir George Villiers of Brooksby, Leicestershire, by his second marriage, to Mary Beaumont, later created Countess of Buckingham. He had two full brothers, George Villiers, successively Viscount Villiers, then Earl, Marquess, and finally Duke of Buckingham, and John Villiers, first Viscount Purbeck, as well as a sister, Susan, later the countess of William Feilding, 1st Earl of Denbigh; while Sir Edward Villiers and Sir George Villiers were his half-brothers. Villiers was described as "unattractive and unintelligent" in his youth, but he shared in his family's good fortune flowing from the position of his brother George as king's favourite. In 1617 he was appointed Gentleman of the Bedchamber to King James I, and on 7 March 1617/18 he was granted an annuity of £200. In December 1617 Sir Robert Naunton a middle-aged man with no sons, was appointed as Secretary of State on the condition of making Villiers his heir, and during his lifetime Villiers gained from Naunton estates yielding £500 a year. Villiers was granted an interest in the monopoly for gold and silver thread, in respect of which he was disappointed to receive only £150 altogether, but he also had a substantial income from the patent for ale houses. In connection with that, he was accused in parliament of malpractice, but the charges were abandoned. In 1620 Villiers had expectations of becoming Master of the Robes, which would secure his position at court. Villiers searched for an heiress to marry, bidding unsuccessfully for the hand of the only daughter of Sir Sebastian Harvey, a rich City of London merchant, and that of Elizabeth Norris, daughter of Francis Norris, 1st Earl of Berkshire and Bridget de Vere. On 16 February 1621/22, a few weeks after Lord Berkshire's suicide on 29 January, John Chamberlain wrote to Sir Dudley Carleton: This heiress was instead won by Edward Wray, another gentleman of the king's bedchamber, and Joseph Mede wrote to Sir Martin Stuteville on 13 April 1622 "Mr. Wray is turned out of the bedchamber for marrying the late Earl of Berkshire's daughter, whom Kit Villiers looked for." On 22 June 1622, Chamberlain wrote to Carleton: "The world talks likewise of divers new earls to be made – as Kit Villiers, if he can be taken off his wench, Earl of Berkshire." Villiers then married Elizabeth, the daughter and heir of Thomas Sheldon of Howley, Leicestershire. On 18 April 1623 Villiers was ennobled as Earl of Anglesey and Baron Villiers of Daventry. This had been in the air for more than a year, with the king seeking to persuade Villiers to give up a mistress, but in the event he had failed to do so. On 19 April, Chamberlain resumed his letters to Carleton: If he had had greater abilities, Villiers might have hoped to gain important positions under the Crown, but as he admitted to his powerful brother in 1627, his "want of preferment proceeded from his own unworthiness rather than from the duke's unwillingness". He was known to have a great fondness for alcohol, and shortly after the death of James I on 27 March 1625 he disappeared from the royal court, with the rumour going about that the new king, Charles I, refused to have him because of his drunkenness. Joseph Mede wrote to Sir Martin Stuteville on 23 April 1625: The appointment of Villiers as Master of the Robes died with the old king, with Charles I retaining Lord Compton, who had served him in that capacity as Prince of Wales. According to one historian of the period, Villiers "summed up all that was unsavory and corrupt with the court of James I". His brother, Buckingham, died on 23 August 1628. Despite this, in December 1628 Villiers gained the position of Keeper of Hampton Court Palace, and in March 1629 he became also Keeper of Bushey Park. He was believed to have bought the reversion of the office of Chancellor of the Exchequer from Edward Barrett, 1st Lord Barrett of Newburgh, but he died at Windsor on 3 April 1630. On 12 April he was entombed there in St George's Chapel. At the time of his death, Villiers was living at Ashley Park, Walton-on-Thames. His widow, Elizabeth, Countess of Anglesey, married secondly Benjamin Weston, Esquire, and they continued to live there. Villiers's titles and estates passed first to his only son, Charles, who died childless on 4 February 1661. Charles had married the widow of his cousin William Villiers, 2nd Viscount Grandison. When she died, many estates passed to Villier's daughter, Anne, widow of Thomas Savile, 1st Earl of Sussex.
